of Species: 'Variation under Domestication' and 'Variation under Nature', and it developed into one of his largest works; Darwin referred to it as his 'big book'. Volume 1 deals with the variations introduced into species as a result of domestication, through changes in climate, diet, breeding and an absence of predators. He began with an examination of dogs and cats, comparing them with their wild counterparts, and moved on to investigate horses and asses; pigs, cattle, sheep, and goats; domestic rabbits; domestic pigeons; fowl; and finally cultivated plants. The work is a masterpiece of nineteenth-century scientific investigation; it is a key text in the development of Darwin's own thought and of the wider discipline of evolutionary biology.

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